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並前往https://www.cwu.edu/primate/sites/cts.cwu.edu.primate/files/Bolt%20et%20al_Ethology_May2019.pdf

Mantled howler monkey (Alouatta palliata) howling behaviour ...

Many primate species have male‐specific long‐distance calls, including platyrrhines like the folivorous howler monkey (Alouatta spp.). Howler monkeys may howl ...

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