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並前往https://books.google.com.tw/books?id=_r4OCAAAQBAJ&pg=RA1-PA401&lpg=RA1-PA401&dq=alleles&source=bl&ots=kwDs0CceWk&sig=ACfU3U28uk8T7LWOxJ_Soz4dHILT51M4UA&hl=zh-TW&sa=X&ved=2ahUKEwjmmsSt67_2AhXlqZUCHepqB70Q6AF6BQjAAhAD

Encyclopedia of Evolutionary Biology - 第 401 頁 - Google 圖書結果

If the allele frequencies are given by pA i and pBj, then D ij 1⁄4 fij À pAipB j (see ... having an arbitrary sign depending on which are the focal alleles, ...

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